In process:
26.  Folk, R.A., R.L. Stubbs, M.E. Mort, N. Cellinese, J.M. Allen, P.S. Soltis, D.E. Soltis, and R.P. Guralnick. In revision. Rates of niche and phenotype evolution lag behind diversification in a temperate radiation. Proceedings of the National Academy of Sciences.

25. Soltis, P.S., R.A. Folk, and D.E. Soltis. In review. Darwin review: Angiosperm phylogeny and evolutionary radiations. Proceedings of the Royal Society B.

24. Sun, M., R.A. Folk, M.A. Gitzendanner, S.A. Smith, C. Germain-Aubrey, R.P. Guralnick, P.S. Soltis, Z. Chen, and D.E. Soltis. Submitted. Exploring the phylogeny and diversification of rosids with a five-locus supermatrix. New Phytologist.
​
23. Liu, L., Y. Du, R.A. Folk, P. Li, D.E. Soltis, and C. Fu. In revision. Plastid genome evolution in Saxifragaceae and multiple plastid capture events involving Heuchera and Tiarella. Molecular Phylogenetics and Evolution.

22. Howard, C.C., R.A. Folk, J.M. Beaulieu, and N. Cellinese. In revision. The monocotyledonous underground: Global climatic and phylogenetic patterns of geophyte diversity. American Journal of Botany.

21. Stubbs, R.L., R.A. Folk, C.-L. Xiang S. Chen, D.E. Soltis, N. Cellinese. Resubmission in prep. Phylogenomics reveals evolutionary processes and historical biogeography within an arctic-alpine lineage (Micranthes, Saxifragaceae). New Phytologist.

20. Yang R., R.A. Folk, N. Zhang, X. Gong. In revision. Formation and population dynamics of hybrids in the Hengduan Mountains flora. Ecology and Evolution.

​Published:
19. Allen, J.M.*, R.A. Folk*, P.S. Soltis, R.P. Guralnick, and D.E. Soltis. 2019. Space, traits, and history: Biodiversity synthesis in the green branches of the Tree of Life. Nature Plants 5: 11–13. https://doi.org/10.1038/s41477-018-0322-7​.

18. Zhang N.-N., Y.-P Ma., R.A. Folk, J.-J. Yu, Y.-Z. Pan, and X. Gong. 2018. Maintenance of species boundaries in three sympatric Ligularia (Senecioneae, Asteraceae) species. Journal of Integrative Plant Biology 60: 986–999. ​Paper: https://doi.org/10.1111/jipb.12674.

17. Allen, J.M., R. LaFrance, R.A. Folk, K. Johnson, R.P. Guralnick. Accepted. aTRAM 2.0: An improved, flexible locus assembler for NGS data. Evolutionary Bioinformatics 14, 1176934318774546. Paper: https://doi.org/10.1177/1176934318774546​.

16. Schuette, S., R.A. Folk, J.T. Cantley, and C.T. Martine. 2018. The hidden Heuchera: How science Twitter uncovered a globally imperiled species in Pennsylvania, USA. PhytoKeys 96: 87-97. Paper: https://doi.org/10.3897/phytokeys.96.23667. 

15. Folk, R.A., C.J. Visger, P.S. Soltis, D.E. Soltis, R.P. Guralnick. Accepted. Geographic range dynamics drove ancient hybridization in a lineage of angiosperms. American Naturalist 192: 171-187. Paper: https://doi.org/10.1086/698120​. Associated Dryad package at: https://doi.org/10.5061/dryad.7cr0c76.

14. Folk, R.A.*, J.C. Ginori*, D.E. Soltis, and A.J. Floden. 2018. Integrative identification of incipient lineages in Heuchera longiflora (Saxifragaceae). Botanical Journal of the Linnean Society 187: 327-345. Paper: https://doi.org/10.1093/botlinnean/boy012. Associated Dryad package at: https://doi.org/10.5061/dryad.6k1s9d4.

13. Stubbs, R.L., R.A. Folk, D.E. Soltis, N. Cellinese. 2018. Pseudo-parallel patterns of disjunctions in an Arctic-alpine plant lineage. Molecular Phylogenetics and Evolution 123: 88-100. Paper: https://www.sciencedirect.com/science/article/pii/S1055790317306760.

12. Folk, R.A., M. Sun, S.A. Smith, P.S. Soltis, R.P. Guralnick, and D.E. Soltis. 2018. Challenges of comprehensive taxon sampling in comparative biology: Wrestling with rosids. Invited review, American Journal of Botany ​105: 433-445. Paper: https://onlinelibrary.wiley.com/doi/full/10.1002/ajb2.1059.

11. Folk, R.A., P.S. Soltis, R.P. Guralnick, and D.E. Soltis, R.P. Guralnick. 2018. New prospects in the detection and comparative analysis of hybridization in the Tree of Life. Invited review, American Journal of Botany 105: 364-375. Paper: https://onlinelibrary.wiley.com/doi/full/10.1002/ajb2.1018

10. Zhang, R., X. Gong, and R.A. Folk. 2017. Evidence for continual hybridization rather than hybrid speciation between Ligularia duciformis and L. paradoxa (Asteraceae). PeerJ 5:e3884. Paper:​ https://peerj.com/articles/3884/.

9. García, N., R.A. Folk, A.W. Meerow, S. Chamala, M.A. Gitzendanner, R.S de Oliveira, D.E. Soltis, and P.S. Soltis. 2017. Deep reticulation and incomplete lineage sorting obscure the diploid phylogeny of rain-lilies and allies (Amaryllidaceae tribe Hippeastreae). Molecular Phylogenetics and Evolution 111: 231-247. Paper: http://www.sciencedirect.com/science/article/pii/S1055790317302865. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.kg686.

8. Freudenstein, J.V., M.B. Broe*, R.A. Folk*, and B.T. Sinn*. 2017. Biodiversity and the species concept – Lineages are not enough. Systematic Biology 66(4): 644–656. Paper: https://academic.oup.com/sysbio/article/66/4/644/2682288.

​7. Folk, R.A., J.R. Mandel, and J.V. Freudenstein. 2017. Ancestral gene flow and parallel organellar genome capture result in extreme phylogenomic discord in a lineage of angiosperms. Systematic Biology 66(3): 320–337. Paper: https://doi.org/10.1093/sysbio/syw083. Associated Dryad package at: ​http://dx.doi.org/10.5061/dryad.cd546.

6. Folk R.A. and P.J. Alexander. 2015. Two new species, Heuchera soltisii and H. inconstans, with further taxonomic notes for the western group of Heuchera section Heuchera (Saxifragaceae). Systematic Botany 40(2): 489-500. Paper: http://www.bioone.org/doi/abs/10.1600/036364415X688853.

5. Folk, R.A., J.R. Mandel, and J.V. Freudenstein. 2015. A protocol for targeted enrichment of intron-containing sequence markers for recent radiations: A phylogenomic example with genomic resources from Heuchera (Saxifragaceae). Applications in Plant Sciences 3(8): 1500039. Paper: http://www.bioone.org/doi/abs/10.3732/apps.1500039. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.4cn66.

4. Folk, R.A. and J.V. Freudenstein. 2015. "Sky islands" in the eastern U.S.A.? – Strong phylogenetic structure in the Heuchera parviflora group (Saxifragaceae). Taxon 64(2): 254-271. Paper: http://www.ingentaconnect.com/content/iapt/tax/2015/00000064/00000002/art00007.

3. Folk, R.A. and J.V. Freudenstein. 2014. Phylogenetic relationships in the genus Heuchera L. (Saxifragaceae) on the basis of nuclear loci. American Journal of Botany 101(9): 1532-1550. Paper: https://doi.org/10.3732/ajb.1400290. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.298d0.

2. Folk, R.A. and J.V. Freudenstein. 2014. Revision of Heuchera section Rhodoheuchera subsections Hemsleyanae and Rosendahliae subsect. nova (Saxifragaceae). Systematic Botany 39(3): 850-874. Paper: http://www.bioone.org/doi/abs/10.1600/036364414X682300.

​1. Folk, R.A. 2013. Heuchera lakelae (Saxifragaceae), a new species from the Sierra La Marta and Sierra Coahuilón, Coahuila and Nuevo León, Mexico. Phytotaxa 124(1): 37-42. Paper: http://www.biotaxa.org/Phytotaxa/article/view/phytotaxa.124.1.4.

*—these authors contributed equally.

In reverse order of acceptance.
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For more information, please visit my Curriculum Vitae.

​I practice open science. My GenBank contributions are here. My Sequence Read Archive contributions can be found by BioProject numbers: PRJNA280966, PRJNA281615, PRJNA374594, PRJNA374874, PRJNA374594.

​I also run a website containing natural history information and extensive photography of all species of Heuchera: https://sites.google.com/site/ryanheuchera/