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Publications

Lab members in BOLD. * means equal authorship. In reverse order of acceptance. Feel free to contact us or use ResearchGate for reprints.

2025
73. Dahal, S., C.M. Siniscalchi, and R.A. Folk. Accepted. A phylogenomic investigation into the biogeography of the Mexico−eastern U.S disjunction in Symphyotrichum (Asteraceae). American Journal of Botany. 

72. Liu, S.-Y.; Y.-Y. Yang, Q. Tian, Z-Y. Yang, H.R. Kates, S.-F. Li, P.J. Valdes, A. Farnsworth, H.R. Kates, C.M. Siniscalchi, R.P. Guralnick, D.E. Soltis, P.S. Soltis, G.W. Stull, R.A. Folk, T.-S. Yi. Accepted. Phylogenomic analyses reveal widespread gene flow during the early radiation of oaks and relatives (Fagaceae: Quercoideae). Journal of Integrative Plant Biology. Preprint available at: https://doi.org/10.1101/2023.04.25.538215

2024
71. Doby, J.R., C.M. Siniscalchi, M. Pajuelo, J. Krigbaum, D.E. Soltis, R.P. Guralnick*, and R.A. Folk*. 2024. Elemental and isotopic analysis of leaves predicts nitrogen-fixing phenotypes. Scientific Reports 14: 20065. https://doi.org/10.1038/s41598-024-70412-8​

​70. Moreno-García, P., Flavia Montaño-Centellas, Y. Liu, E.Y. Reyes-Mendez, R.R. Jha, R.P. Guralnick, R.A. Folk [...; 40 authors in total] D. Li. 2024. Long-term nitrogen deposition reduces the diversity of nitrogen-fixing plants. Science Advances 10: eadp7953. https://doi.org/10.1126/sciadv.adp7953 

69. Gu, W., T. Zhang, S.-Y. Liu, Q. Tian, C.-X. Yang, Q. Lu, X.-G. Fu, H.R. Kates, G.W. Stull, P.S. Soltis, D.E. Soltis, R.A. Folk, R.P. Guralnick, D.-Z. Li, T.-S. Yi. Accepted. Phylogenomics, reticulation, and biogeographical history of Elaeagnaceae. Plant Diversity. https://doi.org/10.1016/j.pld.2024.07.001

68. Pantinople, D., R. Conner, S. Dauber, K. Broussard, N.J. Engle-Wrye, H.R. Jordan, C.M. Siniscalchi, and R.A. Folk. Accepted. Continental sampling reveals core bacterial and environmentally-driven fungal leaf endophytes in Heuchera. American Journal of Botany. Preprint available at: https://doi.org/10.1101/2023.05.23.541975

67. Kates, H.R., B.C. O’Meara, R. LaFrance, G.W. Stull, E.K. James, S. Liu, Q. Tian, T. Yi, J.-M. Ané, D. Conde, M. Kirst, D.E. Soltis, R.P. Guralnick, P.S. Soltis, and R.A. Folk. 2024. Two shifts in evolutionary lability underlie independent gains and losses of root-nodule symbiosis in a single clade of plants. Nature Communications 15: 4262. https://doi.org/10.1038/s41467-024-48036-3

66. Folk, R.A., C.M. Siniscalchi, J. Doby, H.R. Kates, S.R. Manchester, P.S. Soltis, D.E. Soltis, R.P. Guralnick, and M. Belitz. 2024. Spatial phylogenetics of Fagales: Investigating the history of temperate forests. Journal of Biogeography. https://doi.org/10.1111/jbi.14840

65. Jiang, Y.-Z., J.-B. Yang, R.A. Folk, J.-L. Zhao, J. Liu, Z.-S. He, S.-X. Yang,, C.-L. Xiang, and X.-Q. Yu. 2024. Species delimitation of tea plants (Camellia sect. Thea) based on next generation super-barcodes. BMC Plant Biology 24: 181. https://doi.org/10.1186/s12870-024-04882-3

64. Folk, R.A., A.A. Maassoumi, C.M. Siniscalchi, H.R. Kates, D.E. Soltis, P.S. Soltis, M.B. Belitz, and R.P. Guralnick. 2024. Phylogenetic diversity and regionalization in the temperate arid zone. Journal of Systematics and Evolution. https://doi.org/10.1111/jse.13077

63. Li, W., R. Wang, M.-F. Liu, R.A. Folk, B. Xue, R.M.K. Saunders. 2024. Climatic and biogeographic processes underlying the diversification of the pantropical and early divergent angiosperm family Annonaceae. Frontiers in Plant Science 15: 1287171. https://doi.org/10.3389/fpls.2024.1287171

62. Pantinople D.J., N.J. Engle-Wrye, and R.A. Folk. Accepted. Heuchera tuckasegeensis sp. nov. (Saxifragaceae), a new species from western North Carolina. Systematic Botany 49: 37–47. https://doi.org/10.1600/036364424X17110456120677​

61. Folk, R.A., J.L.M. Charboneau, M. Belitz, T. Singh, H.R. Kates, D.E. Soltis, P.S. Soltis, R.P. Guralnick, and C.M. Siniscalchi. 2024. Anatomy of a mega-radiation: Biogeography and niche evolution in Astragalus. American Journal of Botany 111: e16299. https://doi.org/10.1002/ajb2.16299

60. Xu, Z., R.A. Folk, M.A. Gitzendanner, G.-W. Hu, P.S. Soltis, D.E. Soltis, and Q.-F. Wang. 2024. Re-examining the placement of Hydrostachys using a large-scale phylogenetic approach. Taxon 73: 237–248. http://doi.org/10.1002/tax.13122​

59. Carruthers, T., M.S. Moerland, J. Ebersbach, A. Favre, R.A. Folk, J.A. Hawkins, A.N. Muellner-Riehl, M. Röser, D.E. Soltis, N. Tkach, W.J. Baker, J.M. de Vos, and W.L. Eiserhardt. 2024. Rampant upslope biome shifting during late-Cenozoic climate cooling in a diverse alpine plant radiation. Nature Communications 15: 1100. https://doi.org/10.1038/s41467-024-45289-w

58. Tian, Q., G. Stull,; J. Kellermann, D. Medan, F. Nge, L. Shuiyin, H.R. Kates, D.E. Soltis, P.S. Soltis, R.P. Guralnick, R.A. Folk, R. Onstein, and T.-S. Yi. 2024. Rapid in-situ diversification rates in Rhamnaceae explain the parallel evolution of high diversity in temperate biomes from global to local scales. New Phytologist 241: 1851–1865​. https://doi.org/10.1111/nph.19504

57. Folk, R.A., R.P. Guralnick, and R.T. LaFrance. 2024. FloraTraiter: Automated parsing of traits from descriptive biodiversity literature. Applications in Plant Sciences 12: e11563. https://doi.org/10.1002/aps3.11563
​
2023
56 bis. Engle-Wrye, N.J. 2023. Morphometrics and phylogenomics reveal a new species of Heuchera (Saxifragaceae) in northeastern México. Systematic Botany 48: 220–230. [1.101] https://doi.org/10.1600/036364423X16847773873071

56. Wang, C., Z.-Y. Yap; P. Wan, K. Chen, R.A. Folk, D.Z. Damrel, W. Barger, A Diamond, C. Horn, G.P. Landry, T. Samarakoon, S. Harvey, D.R. Morgan, Y. Qiu, and P. Li. 2023. Molecular phylogeography and historical demography of a widespread herbaceous species from eastern North America, Podophyllum peltatum. American Journal of Botany 48: 595-607. https://doi.org/10.1002/ajb2.16254 

​55. Dahal, S., C.M. Siniscalchi, and R.A. Folk. 2023. Species delimitation in the Symphyotrichum subulatum group (Asteraceae) reveals a new species in central Mexico. Systematic Botany 48: 595–607. https://doi.org/10.1600/036364423X17000842213579​ 
​
54. Zhang, Q., R.A. Folk, Z. Mo, H. Ye, Z. Zhang, H. Peng, J. Zhao, S. Yang, X. Yu. 2023. Phylotranscriptomic analyses reveal deep gene tree discordance in Camellia (Theaceae). Molecular Phylogenetics and Evolution. 188: 107912. https://doi.org/10.1016/j.ympev.2023.107912

53. Yang, C.-X., S.-Y. Liu N.J.C. Zerega, G.W. Stull, E.M. Gardner, Q. Tian, W. Gu, Q. Lu, R.A. Folk, H.R. Kates, R.P. Guralnick, D.E. Soltis, P.S. Soltis, Y.-H. Wang, and T. Yi. 2023. Phylogeny and biogeography of Morus (Moraceae). Agronomy 13: 2021. https://doi.org/10.3390/agronomy13082021​

52. Liu, L., M. Chen, R.A. Folk, M. Wang, T. Zhao, F. Shang, D.E. Soltis, and P. Li. Online early. Phylogenomic and syntenic data demonstrate complex evolutionary processes in early radiation of the rosids. Molecular Ecology Resources. https://doi.org/10.1111/1755-0998.13833

51. Folk R.A.*, M.L.Gaynor*, N.J. Engle-Wrye, B.C. O’Meara, P.S. Soltis, D.E. Soltis, R.P. Guralnick, S.A. Smith, C.J. Grady, and Y. Okuyama. 2023. Identifying climatic drivers of hybridization with a new ancestral niche reconstruction method. Systematic Biology 72: 856–873. https://doi.org/10.1093/sysbio/syad018

50. Siniscalchi C.M., J.R. Ackerfield, R.A. Folk. 2023. Diversification and biogeography of North American thistles (Cirsium: Carduoideae: Compositae): Drivers of a rapid continent-wide radiation. International Journal of Plant Sciences 184: 322-341. https://doi.org/10.1086/724310

49. Naranjo A.A., R.A. Folk, M.A. Gitzendanner, D.E. Soltis, P.S. Soltis. 2023. Ancestral area and divergence time analyses reveal Pleistocene-influenced biogeographic evolution in a clade of narrow coastal plain endemic plants. Journal of Biogeography 50: 393-405. https://doi.org/10.1111/jbi.14541

48. Xiao-Gang, F.; S.Y. Liu, R. van Velzen, G. Stull, Q. Tian, R.A. Folk, R.P. Guralnick, H.R. Kates, J.-J. Jin, Z. Li, D.E. Soltis, P.S. Soltis, T.-S. Yi. Online early. Phylogenomic analysis of the hemp family (Cannabaceae) reveals deep cyto-nuclear discordance and provides new insights into generic relationships. Journal of Systematics and Evolution 61: 806-826. https://doi.org/10.1111/jse.12920
​
2022
47. Yu, X.-Q., Y.-Z. Jiang, R.A. Folk, J.-L. Zhao, C.-N. Fu, L. Fang, H. Peng, J.-B. Yang, S.-X. Yang. 2022. Species discrimination in Schima (Theaceae): Next-generation super-barcodes meet evolutionary complexity. Molecular Ecology Resources 22: 3161–3175. https://doi.org/10.1111/1755-0998.13683

46. Doby, J.R., D. Li, R.A. Folk, C.M. Siniscalchi, R.P. Guralnick. 2022. Aridity drives diversity of nitrogen fixing plants in North America. Global Ecology and Biogeography 31: 1630-1642. https://doi.org/10.1111/geb.13535

45. Pereira, W.J., S. Knaack, D. Conde, S. Chakraborty, R.A. Folk, P.M. Triozzi, K.M. Balmant, C. Dervinis, H.W. Schmidt, J.-M. Ané, S. Roy, M. Kirst. 2022. Functional and comparative genomics reveals conserved noncoding sequences in the nitrogen-fixing clade. New Phytologist 234: 634-649​. https://doi.org/10.1111/nph.18006

44. Zhang, Q., L. Zhao, J.-L. Zhao, R.A. Folk, N. Zamora, P.S. Soltis, D.E. Soltis, S.-X. Yang, L.-M. Gao, H. Peng, X.-Q. Yu. 2022. Phylotranscriptomics of Theaceae: generic level relationships, reticulation and whole-genome duplication. Annals of Botany 129: 457–471. https://doi.org/10.1093/aob/mcac007

43. Gorneau, J.A., W.I. Ausich, S. Bertolino, H. Bik, M. Daly, S. Demissew, D.A. Donoso, R.A. Folk, A. Freire-Fierro, S.A. Ghazanfar, O.M. Grace, A.-Q. Hu, S. Kulkarni, I.H. Lichter-Marck, L.G. Lohmann, J. Malumbres-Olarte, A.M. Muasya, A. Pérez-González, Yashica Singh, C.M. Siniscalchi, C.D. Specht, A.L. Stigall, D.C. Tank, L.A. Walker, D.F. Wright, A. Zamani, L.A. Esposito. 2022. Framing the future for taxonomic monography: Improving recognition, support, and access. Bulletin of the Society of Systematic Biologists 1: 8328. https://doi.org/10.18061/bssb.v1i1.8328

42. Figueroa, H., H. Marx, M.B. de Souza Cortez, C.J. Grady, N.J. Engle-Wrye, J. Beach, A. Stewart, R.A. Folk, D.E. Soltis, P.S. Soltis, A. Stewart, and S.A. Smith. 2022. Contrasting patterns of phylogenetic diversity and alpine specialization across the alpine flora of the American mountain range system. Alpine Botany 132: 107–122. https://doi.org/10.1007/s00035-021-00261-y

2021
41. Guo, X., D.-M. Fang, S. Sahu, S. Yang, X. Guang, R.A. Folk, S.A. Smith, A. Chanderbali, S. Chen, M. Liu, T. Yang, S. Zhang, X. Liu, Xun Xu, P.S. Soltis, D.E. Soltis, and H. Liu. 2021. Chloranthus genome provides insights into the early diversification of angiosperms. Nature Communications 12: 6930.

40. Kates, H.R., J.R. Doby, C.M. Siniscalchi, R. Lafrance, D.E. Soltis, P.S. Soltis, R.P. Guralnick, and R.A. Folk. 2021. The effects of herbarium specimen characteristics on short-read NGS sequencing success in nearly 8000 specimens: Old, degraded samples have lower DNA yields but consistent sequencing success. Frontiers in Plant Science 12: 669064. https://doi.org/10.3389/fpls.2021.669064​

39. Folk, R.A., C.M. Siniscalchi. 2021. Biodiversity at the global scale: The synthesis continues. American Journal of Botany 108: 912-924. https://doi.org/10.1002/ajb2.1694​

38. Folk, R.A.*, H.R. Kates*, R. LaFrance, D.E. Soltis, P.S. Soltis, and R.P. Guralnick. 2021. High-throughput methods for efficiently building massive phylogenies from natural history collections. Applications in Plant Sciences 9: e11410. https://doi.org/10.1002/aps3.11410​
​
37. Folk, R.A., R.L. Stubbs, N.J. Engle-Wrye, D.E. Soltis, and Y. Okuyama. 2021. Biogeography and habitat evolution of Saxifragaceae, with a revision of generic limits and a new tribal system. Taxon 70: 263-285. https://doi.org/10.1002/tax.12450​

36. M.B. de Souza Cortez, R.A. Folk, C.J. Grady, J.P. Spoelhof, D.E. Soltis, S.A. Smith, and P.S. Soltis. 2021. Is the age of plant communities predicted by the age, stability, and soil composition of the underlying landscapes?: An investigation of Ocbils. Biological Journal of the Linnean Society ​133: 297–316. https://doi.org/10.1093/biolinnean/blaa174

2020​
35. Folk, R.A., C.M. Siniscalchi, and D.E. Soltis. 2020. Angiosperms at the edge: Extremity, diversity, and phylogeny. Plant, Cell & Environment. https://doi.org/10.1111/pce.13887​ 

34. Stubbs, R.L., R.A. Folk, D.E. Soltis, N. Cellinese. Accepted. Diversification in the arctic: Biogeography and systematics of the North American Micranthes (Saxifragaceae). Systematic Botany 45:802-811. https://doi.org/10.1600/036364420X16033963649282 

33. Folk, R.A.*, N. Sewnath*, C.-L. Xiang, B.T. Sinn, R.P. Guralnick. 2020. Degradation of key photosynthetic genes in the critically endangered semi-aquatic flowering plant Saniculiphyllum guangxiense (Saxifragaceae). BMC Plant Biology 20:324. ​https://doi.org/10.1186/s12870-020-02533-x.

32. Sun, M.*, R.A. Folk*, M.A. Gitzendanner, P.S. Soltis, Z. Chen, D.E. Soltis, and R.P. Guralnick. 2020. Recent, accelerated diversification in rosids occurred outside the tropics. Nature Communications 11: 3333. https://doi.org/10.1038/s41467-020-17116-5.

31. Liu, L., Y. Du, R.A. Folk, P. Li, D.E. Soltis, and C. Fu. 2020. Plastid genome evolution in Saxifragaceae and multiple plastid capture events involving Heuchera and Tiarella. Frontiers in Plant Sciences 11:361​. https://doi.org/10.3389/fpls.2020.00361.
​
30. Sun, M.*, R.A. Folk*, M.A. Gitzendanner, P.S. Soltis, Z. Chen, D.E. Soltis, and R.P. Guralnick. 2020. Estimating rates and patterns of diversification with incomplete sampling: A case study in the rosids. American Journal of Botany 107: 895-909. https://doi.org/10.1002/ajb2.1479.

29. Valle, N., P. Antonenko, P.S. Soltis, D.E. Soltis, R.A. Folk, R.P. Guralnick, J.C. Oliverio, T.T. Difato, Z. Xu, and L. Cheng. 2020. Informal multimedia biodiversity awareness event as a digital ecology for promoting culture of science. Education and Information Technologies 25: 3275–3297​. https://doi.org/10.1007/s10639-020-10121-7.

28. Jantzen J.R., A.P.P.R. Amarasinghe, R.A. Folk, M. Reginato, F.A. Michelangeli, D.E. Soltis, N. Cellinese, P.S. Soltis. 2020. A two-tier bioinformatic pipeline to develop probes for target capture of nuclear loci with applications in Melastomataceae. Applications in Plant Sciences 8: e11345. https://doi.org/10.1002/aps3.11345.

27. Stubbs, R.L., R.A. Folk, C.-L. Xiang S. Chen, D.E. Soltis, N. Cellinese. 2020. A phylogenomic perspective on evolution and discordance in the alpine-Arctic plant clade Micranthes (Saxifragaceae). Frontiers in Plant Sciences 10: 1773. https://doi.org/10.3389/fpls.2019.01773.

26. Oswald, J.A., J.M. Allen, M.J. LeFebvre, B.J. Stucky, R.A. Folk, N.A. Albury, R.P. Guralnick, G.S. Morgan, D.W. Steadman. 2020. Ancient DNA and high-resolution chronometry reveal a long-term human role in the historical diversity and biogeography of the Bahamian hutia. Scientific Reports 10: 1373​. https://doi.org/10.1038/s41598-020-58224-y.

25. Barve, V., L. Brenskelle, D. Li, B. Stucky, N. Barve, M. Hantak, B. McLean, D. Paluh, J. Oswald, M. Belitz, R.A. Folk, R.P. Guralnick. 2020. Methods for broad-scale plant phenology assessments using citizen scientists' photographs. Applications in Plant Sciences 8: e11315. https://doi.org/10.1002/aps3.11315.

2019​
24. Oswald, J.A., J.M. Allen, K.E. Witt, R.A. Folk, N.A. Albury, D.W. Steadman, and R.P. Guralnick. 2019. Ancient DNA from a 2,500-year-old Caribbean fossil places an extinct bird (Caracara creightoni) in a phylogenetic context. Molecular Phylogenetics and Evolution 140: 106576. https://doi.org/10.1016/j.ympev.2019.106576.

23. Yang R., R.A. Folk, N. Zhang, X. Gong. 2019. Homoploid hybridization of plants in the Hengduan mountains region. Ecology and Evolution 9: 8399–8410. https://doi.org/10.1002/ece3.5393.

22.  Folk, R.A., R.L. Stubbs, M.E. Mort, N. Cellinese, J.M. Allen, P.S. Soltis, D.E. Soltis, and R.P. Guralnick. 2019. Rates of niche and phenotype evolution lag behind diversification in a temperate radiation. Proceedings of the National Academy of Sciences 116: 10874–10882. https://doi.org/10.1073/pnas.1817999116. Associated Dryad package at: https://doi.org/10.5061/dryad.cb8gd26

21. Howard, C.C., R.A. Folk, J.M. Beaulieu, and N. Cellinese. 2019. The monocotyledonous underground: Global climatic and phylogenetic patterns of geophyte diversity. American Journal of Botany 106: 850–863. https://doi.org/10.1002/ajb2.1289.

20. Soltis, P.S., R.A. Folk, and D.E. Soltis. 2019. Darwin review: Angiosperm phylogeny and evolutionary radiations. Proceedings of the Royal Society B 286: 20190099. https://doi.org/10.1098/rspb.2019.0099.

19. Allen, J.M.*, R.A. Folk*, P.S. Soltis, R.P. Guralnick, and D.E. Soltis. 2019. Space, traits, and history: Biodiversity synthesis in the green branches of the Tree of Life. Nature Plants 5: 11–13. https://doi.org/10.1038/s41477-018-0322-7​.

2018
18. Zhang N.-N., Y.-P Ma., R.A. Folk, J.-J. Yu, Y.-Z. Pan, and X. Gong. 2018. Maintenance of species boundaries in three sympatric Ligularia (Senecioneae, Asteraceae) species. Journal of Integrative Plant Biology 60: 986–999. ​https://doi.org/10.1111/jipb.12674
​

17. Allen, J.M., R. LaFrance, R.A. Folk, K. Johnson, R.P. Guralnick. Accepted. aTRAM 2.0: An improved, flexible locus assembler for NGS data. Evolutionary Bioinformatics 14, 1176934318774546. https://doi.org/10.1177/1176934318774546

16. Schuette, S., R.A. Folk, J.T. Cantley, and C.T. Martine. 2018. The hidden Heuchera: How science Twitter uncovered a globally imperiled species in Pennsylvania, USA. PhytoKeys 96: 87-97. https://doi.org/10.3897/phytokeys.96.23667

15. Folk, R.A., C.J. Visger, P.S. Soltis, D.E. Soltis, R.P. Guralnick. Accepted. Geographic range dynamics drove ancient hybridization in a lineage of angiosperms. American Naturalist 192: 171-187. Paper: https://doi.org/10.1086/698120​. Associated Dryad package at: https://doi.org/10.5061/dryad.7cr0c76.

14. Folk, R.A.*, J.C. Ginori*, D.E. Soltis, and A.J. Floden. 2018. Integrative identification of incipient lineages in Heuchera longiflora (Saxifragaceae). Botanical Journal of the Linnean Society 187: 327-345. Paper: https://doi.org/10.1093/botlinnean/boy012. Associated Dryad package at: https://doi.org/10.5061/dryad.6k1s9d4.

13. Stubbs, R.L., R.A. Folk, D.E. Soltis, N. Cellinese. 2018. Pseudo-parallel patterns of disjunctions in an Arctic-alpine plant lineage. Molecular Phylogenetics and Evolution 123: 88-100. https://doi.org/10.1016/j.ympev.2018.02.016

12. Folk, R.A., M. Sun, S.A. Smith, P.S. Soltis, R.P. Guralnick, and D.E. Soltis. 2018. Challenges of comprehensive taxon sampling in comparative biology: Wrestling with rosids. American Journal of Botany ​105: 433-445. https://doi.org/10.1002/ajb2.1059

11. Folk, R.A., P.S. Soltis, R.P. Guralnick, and D.E. Soltis, R.P. Guralnick. 2018. New prospects in the detection and comparative analysis of hybridization in the Tree of Life. American Journal of Botany 105: 364-375. https://doi.org/10.1002/ajb2.1018

2017
10. Zhang, R., X. Gong, and R.A. Folk. 2017. Evidence for continual hybridization rather than hybrid speciation between Ligularia duciformis and L. paradoxa (Asteraceae). PeerJ 5:e3884. https://doi.org/10.7717/peerj.3884

9. García, N., R.A. Folk, A.W. Meerow, S. Chamala, M.A. Gitzendanner, R.S de Oliveira, D.E. Soltis, and P.S. Soltis. 2017. Deep reticulation and incomplete lineage sorting obscure the diploid phylogeny of rain-lilies and allies (Amaryllidaceae tribe Hippeastreae). Molecular Phylogenetics and Evolution 111: 231-247. Paper: https://doi.org/10.1016/j.ympev.2017.04.003. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.kg686.

8. Freudenstein, J.V., M.B. Broe*, R.A. Folk*, and B.T. Sinn*. 2017. Biodiversity and the species concept – Lineages are not enough. Systematic Biology 66(4): 644–656. https://doi.org/10.1093/sysbio/syw098

​7. Folk, R.A., J.R. Mandel, and J.V. Freudenstein. 2017. Ancestral gene flow and parallel organellar genome capture result in extreme phylogenomic discord in a lineage of angiosperms. Systematic Biology 66(3): 320–337. Paper: https://doi.org/10.1093/sysbio/syw083. Associated Dryad package at: ​http://dx.doi.org/10.5061/dryad.cd546.

2016
No papers. :(

2015
6. Folk R.A. and P.J. Alexander. 2015. Two new species, Heuchera soltisii and H. inconstans, with further taxonomic notes for the western group of Heuchera section Heuchera (Saxifragaceae). Systematic Botany 40(2): 489-500. Paper: http://www.bioone.org/doi/abs/10.1600/036364415X688853.

5. Folk, R.A., J.R. Mandel, and J.V. Freudenstein. 2015. A protocol for targeted enrichment of intron-containing sequence markers for recent radiations: A phylogenomic example with genomic resources from Heuchera (Saxifragaceae). Applications in Plant Sciences 3(8): 1500039. Paper: http://www.bioone.org/doi/abs/10.3732/apps.1500039. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.4cn66.

4. Folk, R.A. and J.V. Freudenstein. 2015. "Sky islands" in the eastern U.S.A.? – Strong phylogenetic structure in the Heuchera parviflora group (Saxifragaceae). Taxon 64(2): 254-271. Paper: http://www.ingentaconnect.com/content/iapt/tax/2015/00000064/00000002/art00007.

2014
3. Folk, R.A. and J.V. Freudenstein. 2014. Phylogenetic relationships in the genus Heuchera L. (Saxifragaceae) on the basis of nuclear loci. American Journal of Botany 101(9): 1532-1550. Paper: https://doi.org/10.3732/ajb.1400290. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.298d0.

2. Folk, R.A. and J.V. Freudenstein. 2014. Revision of Heuchera section Rhodoheuchera subsections Hemsleyanae and Rosendahliae subsect. nova (Saxifragaceae). Systematic Botany 39(3): 850-874. Paper: http://www.bioone.org/doi/abs/10.1600/036364414X682300.

2013
​1. Folk, R.A. 2013. Heuchera lakelae (Saxifragaceae), a new species from the Sierra La Marta and Sierra Coahuilón, Coahuila and Nuevo León, Mexico. Phytotaxa 124(1): 37-42. Paper: http://www.biotaxa.org/Phytotaxa/article/view/phytotaxa.124.1.4.

PI dissertation: Folk, R.A. 2015. Biosystematics of the Genus Heuchera (Saxifragaceae)​. The Ohio State University. 594 pp.

​We practice open science. Find GenBank contributions here. Sequence Read Archive contributions can be found by BioProject numbers: PRJNA280966, PRJNA281615, PRJNA374594, PRJNA374874, PRJNA374594, PRJNA492276, PRJNA641968. Note also the Dryad links above.

Photo: Hamamelis ×intermedia at NYBG.
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