Lab members in BOLD. * means equal authorship. In reverse order of acceptance. Feel free to contact us or use ResearchGate for reprints.
2023 56. Wang, C., Z.-Y. Yap; P. Wan, K. Chen, R.A. Folk, D.Z. Damrel, W. Barger, A Diamond, C. Horn, G.P. Landry, T. Samarakoon, S. Harvey, D.R. Morgan, Y. Qiu, and P. Li. Molecular phylogeography and historical demography of a widespread herbaceous species from eastern North America, Podophyllum peltatum. American Journal of Botany.
55. Dahal, S., C.M. Siniscalchi, and R.A. Folk. Accepted. Species delimitation in the Symphyotrichum subulatum group (Asteraceae) reveals a new species in central Mexico. Systematic Botany.
54. Zhang, Q., R.A. Folk, Z. Mo, H. Ye, Z. Zhang, H. Peng, J. Zhao, S. Yang, X. Yu. Accepted. Phylotranscriptomic analyses reveal deep gene tree discordance in Camellia (Theaceae). Molecular Phylogenetics and Evolution. https://doi.org/10.1016/j.ympev.2023.107912
53. Yang, C.-X., S.-Y. Liu N.J.C. Zerega, G.W. Stull, E.M. Gardner, Q. Tian, W. Gu, Q. Lu, R.A. Folk, H.R. Kates, R.P. Guralnick, D.E. Soltis, P.S. Soltis, Y.-H. Wang, and T. Yi. 2023. Phylogeny and biogeography of Morus (Moraceae). Agronomy 13: 2021.. https://doi.org/10.3390/agronomy13082021
52. Liu, L., M. Chen, R.A. Folk, M. Wang, T. Zhao, F. Shang, D.E. Soltis, and P. Li. Online early. Phylogenomic and syntenic data demonstrate complex evolutionary processes in early radiation of the rosids. Molecular Ecology Resources. https://doi.org/10.1111/1755-0998.13833
51. Folk R.A.*, M.L.Gaynor*, N.J. Engle-Wrye, B.C. O’Meara, P.S. Soltis, D.E. Soltis, R.P. Guralnick, S.A. Smith, C.J. Grady, and Y. Okuyama. 2023. Identifying climatic drivers of hybridization with a new ancestral niche reconstruction method. Systematic Biology 72: 856–873. https://doi.org/10.1093/sysbio/syad018
50. Siniscalchi C.M., J.R. Ackerfield, R.A. Folk. 2023. Diversification and biogeography of North American thistles (Cirsium: Carduoideae: Compositae): Drivers of a rapid continent-wide radiation. International Journal of Plant Sciences 184: 322-341. https://doi.org/10.1086/724310
49. Naranjo A.A., R.A. Folk, M.A. Gitzendanner, D.E. Soltis, P.S. Soltis. 2023. Ancestral area and divergence time analyses reveal Pleistocene-influenced biogeographic evolution in a clade of narrow coastal plain endemic plants. Journal of Biogeography 50: 393-405. https://doi.org/10.1111/jbi.14541
48. Xiao-Gang, F.; S.Y. Liu, R. van Velzen, G. Stull, Q. Tian, R.A. Folk, R.P. Guralnick, H.R. Kates, J.-J. Jin, Z. Li, D.E. Soltis, P.S. Soltis, T.-S. Yi. Online early. Phylogenomic analysis of the hemp family (Cannabaceae) reveals deep cyto-nuclear discordance and provides new insights into generic relationships. Journal of Systematics and Evolution 61: 806-826. https://doi.org/10.1111/jse.12920 2022 47. Yu, X.-Q., Y.-Z. Jiang, R.A. Folk, J.-L. Zhao, C.-N. Fu, L. Fang, H. Peng, J.-B. Yang, S.-X. Yang. 2022. Species discrimination in Schima (Theaceae): Next-generation super-barcodes meet evolutionary complexity. Molecular Ecology Resources 22: 3161–3175. https://doi.org/10.1111/1755-0998.13683
46. Doby, J.R., D. Li, R.A. Folk, C.M. Siniscalchi, R.P. Guralnick. 2022. Aridity drives diversity of nitrogen fixing plants in North America. Global Ecology and Biogeography 31: 1630-1642. https://doi.org/10.1111/geb.13535
45. Pereira, W.J., S. Knaack, D. Conde, S. Chakraborty, R.A. Folk, P.M. Triozzi, K.M. Balmant, C. Dervinis, H.W. Schmidt, J.-M. Ané, S. Roy, M. Kirst. 2022. Functional and comparative genomics reveals conserved noncoding sequences in the nitrogen-fixing clade. New Phytologist 234: 634-649. https://doi.org/10.1111/nph.18006
44. Zhang, Q., L. Zhao, J.-L. Zhao, R.A. Folk, N. Zamora, P.S. Soltis, D.E. Soltis, S.-X. Yang, L.-M. Gao, H. Peng, X.-Q. Yu. 2022. Phylotranscriptomics of Theaceae: generic level relationships, reticulation and whole-genome duplication. Annals of Botany 129: 457–471. https://doi.org/10.1093/aob/mcac007
43. Gorneau, J.A., W.I. Ausich, S. Bertolino, H. Bik, M. Daly, S. Demissew, D.A. Donoso, R.A. Folk, A. Freire-Fierro, S.A. Ghazanfar, O.M. Grace, A.-Q. Hu, S. Kulkarni, I.H. Lichter-Marck, L.G. Lohmann, J. Malumbres-Olarte, A.M. Muasya, A. Pérez-González, Yashica Singh, C.M. Siniscalchi, C.D. Specht, A.L. Stigall, D.C. Tank, L.A. Walker, D.F. Wright, A. Zamani, L.A. Esposito. 2022. Framing the future for taxonomic monography: Improving recognition, support, and access. Bulletin of the Society of Systematic Biologists 1: 8328. https://doi.org/10.18061/bssb.v1i1.8328
42. Figueroa, H., H. Marx, M.B. de Souza Cortez, C.J. Grady, N.J. Engle-Wrye, J. Beach, A. Stewart, R.A. Folk, D.E. Soltis, P.S. Soltis, A. Stewart, and S.A. Smith. 2022. Contrasting patterns of phylogenetic diversity and alpine specialization across the alpine flora of the American mountain range system. Alpine Botany 132: 107–122. https://doi.org/10.1007/s00035-021-00261-y
2021 41. Guo, X., D.-M. Fang, S. Sahu, S. Yang, X. Guang, R.A. Folk, S.A. Smith, A. Chanderbali, S. Chen, M. Liu, T. Yang, S. Zhang, X. Liu, Xun Xu, P.S. Soltis, D.E. Soltis, and H. Liu. 2021. Chloranthus genome provides insights into the early diversification of angiosperms. Nature Communications 12: 6930.
40. Kates, H.R., J.R. Doby, C.M. Siniscalchi, R. Lafrance, D.E. Soltis, P.S. Soltis, R.P. Guralnick, and R.A. Folk. 2021. The effects of herbarium specimen characteristics on short-read NGS sequencing success in nearly 8000 specimens: Old, degraded samples have lower DNA yields but consistent sequencing success. Frontiers in Plant Science 12: 669064. https://doi.org/10.3389/fpls.2021.669064
39. Folk, R.A., C.M. Siniscalchi. 2021. Biodiversity at the global scale: The synthesis continues. American Journal of Botany 108: 912-924. https://doi.org/10.1002/ajb2.1694
38. Folk, R.A.*, H.R. Kates*, R. LaFrance, D.E. Soltis, P.S. Soltis, and R.P. Guralnick. 2021. High-throughput methods for efficiently building massive phylogenies from natural history collections. Applications in Plant Sciences 9: e11410. https://doi.org/10.1002/aps3.11410 37. Folk, R.A., R.L. Stubbs, N.J. Engle-Wrye, D.E. Soltis, and Y. Okuyama. 2021. Biogeography and habitat evolution of Saxifragaceae, with a revision of generic limits and a new tribal system. Taxon 70: 263-285. https://doi.org/10.1002/tax.12450
36. M.B. de Souza Cortez, R.A. Folk, C.J. Grady, J.P. Spoelhof, D.E. Soltis, S.A. Smith, and P.S. Soltis. 2021. Is the age of plant communities predicted by the age, stability, and soil composition of the underlying landscapes?: An investigation of Ocbils. Biological Journal of the Linnean Society 133: 297–316. https://doi.org/10.1093/biolinnean/blaa174
2020 35. Folk, R.A., C.M. Siniscalchi, and D.E. Soltis. 2020. Angiosperms at the edge: Extremity, diversity, and phylogeny. Plant, Cell & Environment. https://doi.org/10.1111/pce.13887
34. Stubbs, R.L., R.A. Folk, D.E. Soltis, N. Cellinese. Accepted. Diversification in the arctic: Biogeography and systematics of the North American Micranthes (Saxifragaceae). Systematic Botany 45:802-811. https://doi.org/10.1600/036364420X16033963649282
33. Folk, R.A.*, N. Sewnath*, C.-L. Xiang, B.T. Sinn, R.P. Guralnick. 2020. Degradation of key photosynthetic genes in the critically endangered semi-aquatic flowering plant Saniculiphyllum guangxiense (Saxifragaceae). BMC Plant Biology 20:324. https://doi.org/10.1186/s12870-020-02533-x.
32. Sun, M.*, R.A. Folk*, M.A. Gitzendanner, P.S. Soltis, Z. Chen, D.E. Soltis, and R.P. Guralnick. 2020. Recent, accelerated diversification in rosids occurred outside the tropics. Nature Communications 11: 3333. https://doi.org/10.1038/s41467-020-17116-5.
31. Liu, L., Y. Du, R.A. Folk, P. Li, D.E. Soltis, and C. Fu. 2020. Plastid genome evolution in Saxifragaceae and multiple plastid capture events involving Heuchera and Tiarella. Frontiers in Plant Sciences 11:361. https://doi.org/10.3389/fpls.2020.00361. 30. Sun, M.*, R.A. Folk*, M.A. Gitzendanner, P.S. Soltis, Z. Chen, D.E. Soltis, and R.P. Guralnick. 2020. Estimating rates and patterns of diversification with incomplete sampling: A case study in the rosids. American Journal of Botany 107: 895-909. https://doi.org/10.1002/ajb2.1479.
29. Valle, N., P. Antonenko, P.S. Soltis, D.E. Soltis, R.A. Folk, R.P. Guralnick, J.C. Oliverio, T.T. Difato, Z. Xu, and L. Cheng. 2020. Informal multimedia biodiversity awareness event as a digital ecology for promoting culture of science. Education and Information Technologies 25: 3275–3297. https://doi.org/10.1007/s10639-020-10121-7.
28. Jantzen J.R., A.P.P.R. Amarasinghe, R.A. Folk, M. Reginato, F.A. Michelangeli, D.E. Soltis, N. Cellinese, P.S. Soltis. 2020. A two-tier bioinformatic pipeline to develop probes for target capture of nuclear loci with applications in Melastomataceae. Applications in Plant Sciences 8: e11345. https://doi.org/10.1002/aps3.11345.
27. Stubbs, R.L., R.A. Folk, C.-L. Xiang S. Chen, D.E. Soltis, N. Cellinese. 2020. A phylogenomic perspective on evolution and discordance in the alpine-Arctic plant clade Micranthes (Saxifragaceae). Frontiers in Plant Sciences 10: 1773. https://doi.org/10.3389/fpls.2019.01773.
26. Oswald, J.A., J.M. Allen, M.J. LeFebvre, B.J. Stucky, R.A. Folk, N.A. Albury, R.P. Guralnick, G.S. Morgan, D.W. Steadman. 2020. Ancient DNA and high-resolution chronometry reveal a long-term human role in the historical diversity and biogeography of the Bahamian hutia. Scientific Reports 10: 1373. https://doi.org/10.1038/s41598-020-58224-y.
25. Barve, V., L. Brenskelle, D. Li, B. Stucky, N. Barve, M. Hantak, B. McLean, D. Paluh, J. Oswald, M. Belitz, R.A. Folk, R.P. Guralnick. 2020. Methods for broad-scale plant phenology assessments using citizen scientists' photographs. Applications in Plant Sciences 8: e11315. https://doi.org/10.1002/aps3.11315.
2019 24. Oswald, J.A., J.M. Allen, K.E. Witt, R.A. Folk, N.A. Albury, D.W. Steadman, and R.P. Guralnick. 2019. Ancient DNA from a 2,500-year-old Caribbean fossil places an extinct bird (Caracara creightoni) in a phylogenetic context. Molecular Phylogenetics and Evolution 140: 106576. https://doi.org/10.1016/j.ympev.2019.106576.
23. Yang R., R.A. Folk, N. Zhang, X. Gong. 2019. Homoploid hybridization of plants in the Hengduan mountains region. Ecology and Evolution 9: 8399–8410. https://doi.org/10.1002/ece3.5393.
22. Folk, R.A., R.L. Stubbs, M.E. Mort, N. Cellinese, J.M. Allen, P.S. Soltis, D.E. Soltis, and R.P. Guralnick. 2019. Rates of niche and phenotype evolution lag behind diversification in a temperate radiation. Proceedings of the National Academy of Sciences 116: 10874–10882. https://doi.org/10.1073/pnas.1817999116. Associated Dryad package at: https://doi.org/10.5061/dryad.cb8gd26
21. Howard, C.C., R.A. Folk, J.M. Beaulieu, and N. Cellinese. 2019. The monocotyledonous underground: Global climatic and phylogenetic patterns of geophyte diversity. American Journal of Botany 106: 850–863. https://doi.org/10.1002/ajb2.1289.
20. Soltis, P.S., R.A. Folk, and D.E. Soltis. 2019. Darwin review: Angiosperm phylogeny and evolutionary radiations. Proceedings of the Royal Society B 286: 20190099. https://doi.org/10.1098/rspb.2019.0099.
19. Allen, J.M.*, R.A. Folk*, P.S. Soltis, R.P. Guralnick, and D.E. Soltis. 2019. Space, traits, and history: Biodiversity synthesis in the green branches of the Tree of Life. Nature Plants 5: 11–13. https://doi.org/10.1038/s41477-018-0322-7.
2018 18. Zhang N.-N., Y.-P Ma., R.A. Folk, J.-J. Yu, Y.-Z. Pan, and X. Gong. 2018. Maintenance of species boundaries in three sympatric Ligularia (Senecioneae, Asteraceae) species. Journal of Integrative Plant Biology 60: 986–999. https://doi.org/10.1111/jipb.12674 17. Allen, J.M., R. LaFrance, R.A. Folk, K. Johnson, R.P. Guralnick. Accepted. aTRAM 2.0: An improved, flexible locus assembler for NGS data. Evolutionary Bioinformatics 14, 1176934318774546. https://doi.org/10.1177/1176934318774546
16. Schuette, S., R.A. Folk, J.T. Cantley, and C.T. Martine. 2018. The hidden Heuchera: How science Twitter uncovered a globally imperiled species in Pennsylvania, USA.PhytoKeys 96: 87-97. https://doi.org/10.3897/phytokeys.96.23667
15. Folk, R.A., C.J. Visger, P.S. Soltis, D.E. Soltis, R.P. Guralnick. Accepted. Geographic range dynamics drove ancient hybridization in a lineage of angiosperms. American Naturalist 192: 171-187. Paper: https://doi.org/10.1086/698120. Associated Dryad package at: https://doi.org/10.5061/dryad.7cr0c76.
14. Folk, R.A.*, J.C. Ginori*, D.E. Soltis, and A.J. Floden. 2018. Integrative identification of incipient lineages in Heuchera longiflora (Saxifragaceae). Botanical Journal of the Linnean Society 187: 327-345. Paper: https://doi.org/10.1093/botlinnean/boy012. Associated Dryad package at: https://doi.org/10.5061/dryad.6k1s9d4.
13. Stubbs, R.L., R.A. Folk, D.E. Soltis, N. Cellinese. 2018. Pseudo-parallel patterns of disjunctions in an Arctic-alpine plant lineage. Molecular Phylogenetics and Evolution 123: 88-100. https://doi.org/10.1016/j.ympev.2018.02.016
12. Folk, R.A., M. Sun, S.A. Smith, P.S. Soltis, R.P. Guralnick, and D.E. Soltis. 2018. Challenges of comprehensive taxon sampling in comparative biology: Wrestling with rosids. American Journal of Botany 105: 433-445. https://doi.org/10.1002/ajb2.1059
11. Folk, R.A., P.S. Soltis, R.P. Guralnick, and D.E. Soltis, R.P. Guralnick. 2018. New prospects in the detection and comparative analysis of hybridization in the Tree of Life. American Journal of Botany 105: 364-375. https://doi.org/10.1002/ajb2.1018
2017 10. Zhang, R., X. Gong, and R.A. Folk. 2017. Evidence for continual hybridization rather than hybrid speciation between Ligularia duciformis and L. paradoxa (Asteraceae). PeerJ 5:e3884. https://doi.org/10.7717/peerj.3884
9. García, N., R.A. Folk, A.W. Meerow, S. Chamala, M.A. Gitzendanner, R.S de Oliveira, D.E. Soltis, and P.S. Soltis. 2017. Deep reticulation and incomplete lineage sorting obscure the diploid phylogeny of rain-lilies and allies (Amaryllidaceae tribe Hippeastreae). Molecular Phylogenetics and Evolution 111: 231-247. Paper: https://doi.org/10.1016/j.ympev.2017.04.003. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.kg686.
8. Freudenstein, J.V., M.B. Broe*, R.A. Folk*, and B.T. Sinn*. 2017. Biodiversity and the species concept – Lineages are not enough. Systematic Biology 66(4): 644–656. https://doi.org/10.1093/sysbio/syw098
7. Folk, R.A., J.R. Mandel, and J.V. Freudenstein. 2017. Ancestral gene flow and parallel organellar genome capture result in extreme phylogenomic discord in a lineage of angiosperms. Systematic Biology 66(3): 320–337. Paper: https://doi.org/10.1093/sysbio/syw083. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.cd546.
2016 No papers. :(
2015 6. Folk R.A. and P.J. Alexander. 2015. Two new species, Heuchera soltisii and H. inconstans, with further taxonomic notes for the western group of Heuchera section Heuchera (Saxifragaceae). Systematic Botany 40(2): 489-500. Paper: http://www.bioone.org/doi/abs/10.1600/036364415X688853.
5. Folk, R.A., J.R. Mandel, and J.V. Freudenstein. 2015. A protocol for targeted enrichment of intron-containing sequence markers for recent radiations: A phylogenomic example with genomic resources from Heuchera (Saxifragaceae). Applications in Plant Sciences 3(8): 1500039. Paper: http://www.bioone.org/doi/abs/10.3732/apps.1500039. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.4cn66.
2014 3. Folk, R.A. and J.V. Freudenstein. 2014. Phylogenetic relationships in the genus Heuchera L. (Saxifragaceae) on the basis of nuclear loci. American Journal of Botany 101(9): 1532-1550. Paper: https://doi.org/10.3732/ajb.1400290. Associated Dryad package at: http://dx.doi.org/10.5061/dryad.298d0.
2. Folk, R.A. and J.V. Freudenstein. 2014. Revision of Heuchera section Rhodoheuchera subsections Hemsleyanae and Rosendahliae subsect. nova (Saxifragaceae). Systematic Botany 39(3): 850-874. Paper: http://www.bioone.org/doi/abs/10.1600/036364414X682300.
2013 1. Folk, R.A. 2013. Heuchera lakelae (Saxifragaceae), a new species from the Sierra La Marta and Sierra Coahuilón, Coahuila and Nuevo León, Mexico. Phytotaxa 124(1): 37-42. Paper: http://www.biotaxa.org/Phytotaxa/article/view/phytotaxa.124.1.4.
PI dissertation: Folk, R.A. 2015. Biosystematics of the Genus Heuchera (Saxifragaceae). The Ohio State University. 594 pp.